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Energy flows through ecosystems while chemical elements can be recycled. Use clearly defined budgets and units to compare production at different trophic levels.
What you'll be able to do
Gross primary production represents the chemical energy fixed into organic compounds before respiratory losses are deducted over the stated area, volume and period. Net primary production is what remains after producers’ respiration: NPP=GPP−R. It can support growth and reproduction and become available to consumers or decomposers.
Biomass may be measured as dry mass or carbon mass per area. Drying reduces the confounding effect of variable water content; calorimetry estimates chemical energy released from biomass combustion under a defined method. Biomass and energy are not interchangeable numerical units, so use any conversion information supplied.
For a consumer, net production can be modelled as N=I−(F+R), where I is the energy in ingested material, F is energy lost in faeces and urine under the question’s definition, and R is respiratory loss. Not all food encountered is eaten, and not all ingested food is assimilated.
Respiration transfers energy to the environment as heat rather than destroying energy. Uneaten biomass and waste can support decomposers, so an energy pathway that does not reach the next named consumer is not necessarily absent from the ecosystem. Productivity is the rate of production, requiring a time unit such as kJ m⁻² year⁻¹.
An efficiency calculation depends on which transfer is being assessed: net production as a percentage of ingestion is different from energy passed between two trophic levels. Label the numerator and denominator before substituting. Real efficiencies vary; there is no universal requirement that each transfer is exactly 10%.
Farming can simplify food webs to direct more production towards human food, and some practices reduce livestock respiratory losses associated with movement or temperature regulation. Evaluate welfare, biodiversity, resource inputs and sustainability alongside yield. A higher transfer efficiency in one narrow budget does not by itself prove lower total environmental impact.
Try each question before opening the worked response. Use precise biological terms, connect each cause to its effect, and check units and assumptions when interpreting data.
Equation recap
Common mistakes to avoid
Key takeaways
Test yourself
Answer a few questions, then review anything you missed.
Start with the quiz above to check the key ideas. You can also choose another topic from the practice library.